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0.15294 0.15294 scn 5.5244 0 TD [(\),)-383.9(and)-379.2(for)-382(this)-382.4(reason)-376.4(legislative)-387.2(measures)-382.6(are)]TJ -8.0378 -1.3921 TD [(coming)-288.6(into)-286.8(place)-291.7(worldwide)-288.4(requiring)-294.4(the)-288(welfare)-286(of)-286.2(these)-289.1(species)]TJ 0 -1.3858 TD 0.016 Tc [(to)-517.1(be)-512.9(accommodated)-516(i)-1.2(n)-520.2(t)3(heir)-520.8(capture,)-515.6(holding)-515(a)-0.5(nd)-516(slaug)-4.7(h)5.3(ter)]TJ 0 -1.3921 TD 0 Tc [(proced)-10.4(ures.)-511.7(Countrie)-12.6(s)-499(tha)-8.7(t)-498(curre)-11.3(ntly)-502.5(pla)-8.7(ce)-503.4(decapo)-13(d)-500.5(crusta)-9.6(ceans)]TJ 0 -1.3858 TD [(under)-385.4(some)-389.9(degree)-389.5(of)-386.9(legal)-381.1(protection)-389.2(include)-390.4(Australia,)-387.7(Austria,)]TJ 0 -1.3921 TD 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[(DEFRA,)-386.5(2022)]TJ 0.15294 0.15294 0.15294 scn 5.6315 0 TD (\))Tj -26.4441 -1.3921 TD [(which)-255.8(recognises)-260(that)-254.8(all)-259.8(decapod)-261.5(crustaceans)-260.8(are)-257(sentient,)-255.7(and)-259.5(this)]TJ 28.3654 52.7244 TD [(Act)-321(will)-320.5(provide)-319.5(oversight)-328.1(of)-317.7(any)-321.5(other)-324.8(new)-318.7(legislation.)-329.8(Calls)-317(have)]TJ 0 -1.3921 TD [(now)-423.7(been)-432.5(made)-429.2(for)-426.1(the)-432.9(use)-426(of)-431(decapod)-431.6(crustaceans)-430.8(in)-433.6(scienti)]TJ /F7 1 Tf 25.8331 0 TD ()Tj /F6 1 Tf 0.5354 0 TD (c)Tj -26.3685 -1.3858 TD [(research)-219.5(to)-218.5(be)-217.9(subject)-224.1(to)-218.5(ethical)-222.6(review)-218.4(\()]TJ 0.30196 0.72157 0.33333 scn 15.6787 0 TD [(Rowe,)-220.6(2018)]TJ 0.15294 0.15294 0.15294 scn 4.5669 0 TD [(\),)-220.1(and)-221.7(also)-219.1(in)-219.5(the)]TJ -20.2457 -1.3921 TD 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1 Tf 0.5291 0 TD [(nition)-348.7(pertaining)-348.2(to)-344.5(the)-344.7(ability)-348.7(of)-342.9(the)-344.7(animal)-349.6(to)]TJ -7.4142 -1.3858 TD [(sense)-280.1(presented)-285.8(stimuli)-280.8(and)-284.7(respond)-283.6(to)-275.2(them,)-284.8(which)-281(directly)-283.3(aligns)]TJ 0 -1.3921 TD [(with)-296.1(the)-300.6(aims)-306.5(of)-298.8(this)-300.5(work.)]TJ 1.5496 -1.3858 TD 0.0217 Tc [(From)-520.6(an)-514.5(animal)-515.7(welfa)5.2(r)1.4(e)-519.8(p)0(erspecti)4.5(v)1(e,)-516.2(bo)5.2(th)-524.5(th)4.7(e)-519.8(o)-1.1(nset)-514.1(and)]TJ -1.5496 -1.3921 TD 0 Tc [(persistence)-402.1(of)-399.5(insensibility)-408.2(are)-401.9(important)-402.9(in)-402.1(order)-399.8(to)-401.1(ensure)-402.6(that)]TJ 0 -1.3858 TD 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[(pr)-10.5(ovid)-13.2(es)-505(im)-11.5(por)-8.1(tan)-11.3(t)-504.3(inf)-10.4(orm)-12.2(ati)-8.9(on)-508.6(on)-508.6(t)-6.7(h)0(e)-508.1(meta)-10.9(bo)-7.9(lic)-510.2(ch)-10.5(ange)-12.5(s)-505.3(a)0(n)-11(d)]TJ 0 -1.3921 TD [(res)-8.7(pons)-8.8(es)-511.3(to)-508.2(env)-8.6(iron)-10.6(menta)-11.7(l)-505.9(pert)-10.6(urb)-8.1(ation)-13.5(s)-505.3(\()]TJ 0.30196 0.72157 0.33333 scn 18.3307 0 TD [(Mc)-9.2(Gaw)-508.7(and)-511.4(Reib)-11.6(er,)]TJ -18.3307 -1.3858 TD (2015)Tj 0.15294 0.15294 0.15294 scn 1.915 0 TD (;)Tj 0.30196 0.72157 0.33333 scn 0.5606 0 TD [(McLean)-345.6(and)-347.7(Todgham,)-344.6(2015)]TJ 0.15294 0.15294 0.15294 scn 11.7669 0 TD (;)Tj 0.30196 0.72157 0.33333 scn 0.5606 0 TD [(Kushinsky)-348.7(et)-340.3(al.,)-344.7(2019)]TJ 0.15294 0.15294 0.15294 scn 8.9827 0 TD [(\),)-346.1(acute)]TJ -23.7858 -1.3921 TD [(stress)-471.6(responses)-472.2(\()]TJ 0.30196 0.72157 0.33333 scn 7.3071 0 TD [(Burnovicz)-471.1(et)-472.5(al.,)-464.4(2009)]TJ 0.15294 0.15294 0.15294 scn 9.2283 0 TD (;)Tj 0.30196 0.72157 0.33333 scn 0.6866 0 TD [(Yazawa,)-469.5(2015)]TJ 0.15294 0.15294 0.15294 scn 5.6315 0 TD [(\))-469.6(and)-467.3(for)]TJ -22.8535 -1.3858 TD [(ecotoxicology)-338.6(studies)-340.6(\()]TJ 0.30196 0.72157 0.33333 scn 9.2094 0 TD [(Bamber)-339.9(and)-335.1(Depledge,)-340.4(1997)]TJ 0.15294 0.15294 0.15294 scn 11.4835 0 TD (;)Tj 0.30196 0.72157 0.33333 scn 0.5543 0 TD (Kholodkevich)Tj -21.2472 -1.3921 TD 0.0123 Tc [(et)-510.9(al.,)-506.7(2017)]TJ 0.15294 0.15294 0.15294 scn 4.8945 0 TD 0.0117 Tc [(\).)-513.6(Monitor)4(i)0.8(ng)-513(HR)-514.5(is)-512.5(also)-508.8(us)4(ed)-514(widely)-506.7(to)-508.8(establish)]TJ -4.8945 -1.3858 TD 0 Tc [(metabolic)-327.4(activity,)-328.3(locomotory)-323.9(performance,)-330.1(agonistic)-329.7(interactions)]TJ T* [(and)-479.9(responses)-484.8(to)-476.7(environmental)-486.4(conditions)-482.6(and)-479.9(biotic)-476.9(variables)]TJ 0 -1.3921 TD (\()Tj 0.30196 0.72157 0.33333 scn 0.3465 0 TD [(Han)-13.2(dy)-508.6(and)-511.4(De)-7.7(ple)-10.8(dge)-10.7(,)-506.4(1999)]TJ 0.15294 0.15294 0.15294 scn 11.6535 0 TD (;)Tj 0.30196 0.72157 0.33333 scn 0.7244 0 TD [(Gre)-11.2(en,)-513.2(2011)]TJ 0.15294 0.15294 0.15294 scn 5.115 0 TD [(\).)-509.9(We)-505.8(ha)-8.4(ve)-511.8(ther)-12.2(efo)-8.7(re)]TJ -17.8394 -1.3858 TD [(assessed)-440(the)-439.2(potential)-440.5(to)-432.6(use)-438.6(HR)-440.2(as)-433.6(an)-439.3(indicator)-444.1(of)-431(the)-439.2(state)-440.4(of)]TJ 0 -1.3921 TD [(sensibi)-9.9(lity)-500.1(in)-502.9(decapod)-507.2(crust)-11.9(aceans)-507.1(by)-497.1(monit)-10.2(oring)-503.8(it)-502.6(in)-502.9(both)]TJ /F9 1 Tf 25.9339 0 TD (C.)Tj -25.9339 -1.3858 TD (maenas)Tj /F6 1 Tf 3.3197 0 TD (and)Tj /F9 1 Tf 1.8016 0 TD [(H.)-298.1(gammarus)]TJ /F6 1 Tf 5.7575 0 TD [(before)-303.5(and)-297.3(after)-306.5(electrical)-302.9(stunning.)]TJ /F1 1 Tf 13.5 0 0 13.5 49.6063 246.3874 Tm 0.30196 0.72157 0.33333 scn [(2)-298.9(Methods)]TJ /F6 1 Tf 9 0 0 9 63.6094 221.3858 Tm 0.15294 0.15294 0.15294 scn [(The)-228(methodologies)-230.8(used)-227.2(to)-224.8(electrically)-231.1(stun)-232(the)]TJ /F9 1 Tf 18.8221 0 TD [(C.)-223.2(maenas)]TJ /F6 1 Tf 4.3276 0 TD (crabs)Tj -24.7055 -1.3858 TD 0.0427 Tc [(and)-539.7(t)-1.8(he)]TJ /F9 1 Tf 4.0882 0 TD 0.042 Tc [(H.)-541.6(gammarus)]TJ /F6 1 Tf 6.6583 0 TD 0.0415 Tc [(lobsters,)-546.8(a)-0.2(nd)-547.2(to)-548.3(make)-544.1(neurologica)-6.5(l)]TJ -10.7465 -1.3921 TD 0 Tc [(recordings,)-416.8(are)-408.2(here)-408.4(summarised)-416(and)-410.7(were)-407.3(the)-407.7(same)-415.1(as)-408.4(reported)]TJ 0 -1.3858 TD (by)Tj 0.30196 0.72157 0.33333 scn 1.2472 0 TD [(Albalat)-292.9(et)-296.2(al.)-291.8(\(2022\))]TJ 0.15294 0.15294 0.15294 scn 8.2646 0 TD (and)Tj 0.30196 0.72157 0.33333 scn 1.789 0 TD [(Neil)-294(et)-289.9(al.)-291.8(\(2022\))]TJ 0.15294 0.15294 0.15294 scn 6.7717 0 TD [(,)-292.2(where)-292(more)-289.1(detailed)]TJ -18.0724 -1.3921 TD [(descriptions)-306.8(can)-300.5(be)-299.7(found.)]TJ /F1 1 Tf 11.9999 0 0 11.9999 49.6063 133.9087 Tm [(2.1)-301.1(Animal)-301.1(supply)-307(and)-301.9(holding)]TJ /F6 1 Tf 9 0 0 9 63.6094 108.9071 Tm (The)Tj /F9 1 Tf 1.978 0 TD [(Carcinus)-419(maenas)]TJ /F6 1 Tf 7.3764 0 TD [(\(L.\))-417(crabs)-419.8(used)-416.2(in)-414.7(these)-421.4(trials)-416.7(were)-419.9(all)]TJ -10.9102 -1.3921 TD [(males)-410.7(of)-405.8(65-75)-407.1(mm)-404.3(carapace)-412.1(width)-406(in)-408.4(the)-407.7(intermoult)-407(stage,)-406.6(and)]TJ 28.3654 73.6126 TD [(were)-413.6(obtained)-423.2(live)-417.6(from)-420.4(commercial)-419.3(suppliers)-428.3(\(UMBSM)-415.6(Animal)]TJ 0 -1.3858 TD [(Supply)-287.6(Service)-294.9(Millport)-297.3(and)-291(Loch)-289.7(Fyne)-294.9(Sea)-291.6(Farms)-293.5(Ltd.,)-289.7(UK\).)-292.8(They)]TJ 0 -1.3921 TD [(were)-300.2(housed)-298.6(initially)-301.7(in)-301.4(communal)-304.1(tanks,)-300.9(within)-301.4(a)-300(closed)-305.6(seawater)]TJ 0 -1.3858 TD [(circulating)-235.8(system)-232(at)-231.1(10C,)-228.7(then)-238.3(retained)-233.7(individually)-236(in)-232.1(tanks)-229.2(for)-237.1(at)]TJ 0 -1.3921 TD [(least)-372.7(two)-379.3(weeks)-378.1(before)-379.1(experimentation.)-389.1(From)-374.1(this)-382.4(stock,)-375.5(6)-377.7(crabs)]TJ 0 -1.3858 TD 0.0212 Tc [(were)-526.6(subjected)-523.4(t)1.9(o)-524.5(e)2.5(lect)-4.4(rical)-522.5(s)0.9(t)-4.4(u)]TJ 13.789 0 TD 0.0214 Tc [(nning)-522.2(for)-521.7(e)-3.6(lec)-3.5(t)2.1(rophysiological)]TJ -13.789 -1.3921 TD 0 Tc [(recording,)-472.1(4)-465.9(crabs)-470.2(were)-470.3(used)-472.9(for)-470.2(measuring)-470.5(HR)-471.7(and)-467.3(another)-474.3(6)]TJ 0 -1.3858 TD [(were)-300.2(used)-302.8(as)-301.4(controls.)]TJ 1.5496 -1.3921 TD [(The)-253.2(European)-247.1(lobsters)-253.4(\()]TJ /F9 1 Tf 9.5748 0 TD [(Homarus)-250.9(gammarus)]TJ /F6 1 Tf 8.3905 0 TD [(\(L.\))-246.9(were)-249.8(all)-247.2(males)]TJ -19.515 -1.3858 TD [(of)-368.1(carapace)-380.6(length)-374.4(80-95)-375.6(mm)-372.8(in)-376.9(the)-376.2(intermoult)-375.5(stage.)-375.1(They)-374.7(were)]TJ 0 -1.3921 TD [(captured)-341.2(by)-339.6(commercial)]TJ /F7 1 Tf 10.1543 0 TD ()Tj /F6 1 Tf 0.5354 0 TD [(shermen)-341.8(using)-338.9(baited)-342.9(traps)-337.4(\(creels\))-344.7(laid)]TJ -10.6898 -1.3858 TD [(of)66.6(fshore)-293(from)-294.4(St)-288.3(Abbs)-292.3(on)-294.4(the)-288(east)-295.2(coast)-292.8(of)-286.2(Scotland.)-297.6(After)-289.6(banding)]TJ 0 -1.3921 TD [(the)-388.8(claws)-388.3(of)-393.2(the)-388.8(lobsters)-391.9(\(a)-391.6(standard)-389(practice)-396.5(by)]TJ /F7 1 Tf 20.3654 0 TD ()Tj /F6 1 Tf 0.5291 0 TD [(shermen\))-395.6(they)]TJ -20.8945 -1.3858 TD [(were)-275(held)-286.1(initially)-276.5(in)-282.5(seawater)-284.6(tanks)-279.6(at)-281.5(the)-275.4(St)-282(Abbs)-279.7(Marine)-284.4(Station,)]TJ 0 -1.3921 TD [(then)-263.5(transferred)-276.1(in)-269.9(chilled)-267.2(containers)-268.4(to)-268.9(the)-269.1(University)-270(of)-267.3(Glasgow.)]TJ 0 -1.3858 TD [(As)-347.4(with)-352.8(the)-351(crabs,)-352.9(they)-352.8(were)-350.6(housed)-355.3(initially)-352.1(in)-351.7(communal)-354.5(tanks,)]TJ 0 -1.3921 TD [(within)-332.9(a)-331.5(closed)-330.8(seawater)-335(circulating)-336.6(system)-339.1(at)-331.9(10C,)-329.5(then)-332.8(retained)]TJ 0 -1.3858 TD [(individually)-261.2(in)-257.3(tanks)-260.7(for)-262.3(at)-262.6(least)-259.3(two)-259.6(weeks)-258.5(before)-265.7(experimentation.)]TJ 0 -1.3921 TD [(From)-267(this)-269(stock,)-274.8(6)-270.6(lobsters)-272.3(were)-268.7(subjected)-277.1(to)-268.9(electrical)-271.4(stunning)-271.4(for)]TJ 0 -1.3858 TD [(electrophysiological)-327.2(recording,)-333.5(6)-327.3(lobsters)-322.7(were)-331.7(used)-328(for)-325.3(measuring)]TJ 0 -1.3921 TD [(HR)-295.3(and)-303.6(another)-304.2(6)-302.1(used)-302.8(as)-301.4(controls.)]TJ /F1 1 Tf 11.9999 0 0 11.9999 304.8944 471.4015 Tm [(2.2)-298.5(Electrical)-302.6(stunning)]TJ /F6 1 Tf 9 0 0 9 318.8409 446.3999 Tm [(Electrical)-390(stunning)-384.8(was)-386.6(applied)-390.4(to)-382.2(individuals)-384.8(of)-386.9(each)-385.8(species)]TJ -1.5496 -1.3921 TD [(without)-460.7(prior)-467.6(anaesthesia)-471.3(using)-471.2(a)-463.8(commercially)-473.5(available)-466.8(device)]TJ 0 -1.3858 TD 0.029 Tc [(\(Crusta)-6.4(s)2.4(tun)]TJ /F10 1 Tf 5.0394 0.3339 TD 0 Tc ()Tj /F6 1 Tf 1.0079 -0.3339 TD 0.0289 Tc [(;)-527.9(S)-0.7(tudham)-532.3(Technol)-4.6(o)-0.2(gies)-526.8(Ltd,)-534.2(Ca)-6.5(mbridge,)-534.2(UK\))]TJ -6.0472 -1.3921 TD 0 Tc [(following)-386.3(the)-388.8(manufacturer)]TJ /F8 1 Tf 11.1811 0 TD ()Tj /F6 1 Tf 0.2142 0 TD [(s)-391.9(operating)-390.9(procedures.)-396.1(The)-391.7(chamber)]TJ -11.3953 -1.3858 TD [(of)-324(the)-325.8(stunning)-328.1(device)-329.7(was)]TJ /F7 1 Tf 11.1685 0 TD ()Tj /F6 1 Tf 0.5291 0 TD [(lled)-327.8(with)-327.6(a)-325.2(salt)-328.9(solution)-328.3(\(NaCl)-327.7(3)-327.3(g/L\))]TJ -11.6976 -1.3921 TD [(and)-259.5(a)-262.2(stunning)-265.1(cycle)-267.6(of)-261(110)-263.8(V,)-262.9(2)]TJ /F8 1 Tf 12.8819 0 TD ()Tj /F6 1 Tf 0.5102 0 TD [(5)-258(amp)-265.6(electrical)-265.1(charge)-263(for)-262.3(5)-264.3(s)-259.6(for)]TJ -13.3921 -1.3858 TD (the)Tj /F9 1 Tf 1.5685 0 TD [(C.)-317.6(maenas)]TJ /F6 1 Tf 4.5102 0 TD [(crabs)-325.3(and)-322.5(10)-320.7(s)-322.6(for)-319(the)-319.5(larger)]TJ /F9 1 Tf 11.8488 0 TD [(H.)-317(gammarus)]TJ /F6 1 Tf 5.7953 0 TD (lobsters)Tj -23.7228 -1.3921 TD [(was)-298.4(applied.)]TJ /F1 1 Tf 11.9999 0 0 11.9999 304.8944 321.3921 Tm [(2.3)-298.5(Animal)-301.1(preparation)]TJ /F6 1 Tf 9 0 0 9 318.8409 296.3905 Tm [(Crabs)-236.5(not)-231.8(to)-231.1(be)-230.4(subjected)-239.3(to)-231.1(electrical)-233.6(stunning)-233.6(were)-237.2(held)-229.5(on)-231.4(ice)]TJ -1.5496 -1.3858 TD [(for)-495.4(30)-490.8(min)-497.2(prior)-499.1(to)-495.6(experimentation)-500.1(to)-495.6(reduce)-497.6(their)-501.8(metabolic)]TJ 0 -1.3921 TD [(activity)-426.7(and)-429.5(to)-426.3(induce)-432.5(torpor,)-432(while)-429.8(those)-431.9(subjected)-428.3(to)-426.3(electrical)]TJ 0 -1.3858 TD [(stunning)-302.9(were)-300.2(used)-302.8(directly)-302.2(after)-306.5(the)-300.6(stun.)]TJ 1.5496 -1.3921 TD [(In)-234.7(order)-229.7(to)-231.1(expose)-232.5(the)-231.3(central)-228.3(nervous)-238.5(system)-232(\(CNS\))-228.9(of)-229.5(the)-231.3(crab)]TJ -1.5496 -1.3858 TD 0.0177 Tc [(f)6.3(o)1.2(r)-519.1(r)3.7(ecordin)4.4(g)-519.6(\()]TJ 0.30196 0.72157 0.33333 scn 6.589 0 TD 0.0152 Tc [(Fig)-5.5(u)-0.2(re)-520(1)]TJ 0.15294 0.15294 0.15294 scn 3.6535 0 TD 0.0171 Tc [(\),)-514.5(the)-518.1(c)-1.5(arapa)-5.7(c)4.8(e)-518.1(w)1.2(as)-519.7(removed)-514.9(a)0.6(nd)-514.9(the)]TJ -10.2425 -1.3921 TD 0.0657 Tc [(prep)-6.4(a)5.1(r)-5(atio)5.1(n)-571.3(w)-0.6(a)5.1(s)-571.9(submerged)-567.1(i)4.4(n)-571.3(a)-561.9(balanced)-567.1(s)-5(a)5.1(l)-5.6(t)-564.6(s)1.4(olution)]TJ 0 -1.3858 TD 0.0795 Tc [(corresp)-5.2(o)0(nding)-577.5(i)-0.7(n)-582.7(c)4.2(omposition)-576.4(a)-6.3(n)3.2(d)-578.5(o)0(smolarity)-577.5(t)3.5(o)-579.6(c)-2.1(ra)-6.3(b)]TJ 0 -1.3921 TD 0 Tc [(haemolymph,)-341.7(at)-344.5(a)-337.8(temperature)-346.7(of)-342.9(10C.)-342.1(The)-341.3(internal)-347.2(organs)-340.7(were)]TJ 0 -1.3858 TD 0.0481 Tc [(t)3.6(h)5.9(en)-538.5(displa)6.4(ce)4.3(d)-546.9(i)5.7(n)-544.8(o)6.4(rd)7.5(er)-539.1(to)-541.7(e)4.3(x)-0.4(po)6.4(se)-543.8(t)3.6(h)-0.4(e)-543.8(c)4.3(irc)4.3(u)1.2(m)3.4(o)0.1(e)4.3(s)2.7(oph)5.9(a)0.1(ge)4.3(al)]TJ 0 -1.3921 TD 0 Tc [(connectives)-302.2(around)-305.4(the)-300.6(base)-298.7(of)-305.1(the)-300.6(stomach.)]TJ 1.5496 -1.3858 TD [(A)-270.5(similar)-270.3(procedure)-272.6(was)-266.9(employed)-271.7(for)-268.6(the)-269.1(lobster,)-267.1(but)-265.4(after)-275(the)]TJ 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[(H)-14.1(a)5(r)-14.1(r)8.4(i)-14.1(n)5.8(g)-14.1(t)8.9(o)-14.1(n)-201.4(e)-14.1(t)-203.2(a)-14.1(l)5.2(.,)-218.1(2020)]TJ 0.15294 0.15294 0.15294 scn 8.6992 0 TD 0 Tc [(\).)-188.6(A)13(b)0(o)23.6(u)0(t)-192.2(3)-195(h)-199.7(p)0(r)21(i)14.3(or)-188(to)-187(ex)21(pe)22.6(ri)19.3(me)18.6(nt)24.1(at)20.9(io)23(n)-202.3(t)0(h)26.7(e)]TJ -10.9858 -1.3921 TD [(an)20.5(i)14.3(m)0(a)20.7(l)-216.1(wa)17.9(s)-209.2(h)0(e)21(l)0(d)-206.4(o)0(n)-206.2(i)14.3(ce)-207.4(f)13.8(o)0(r)-206.9(3)12.9(0)-213.9(m)0(i)20(n)-214.9(to)-205.9(re)24.1(du)19.5(c)12.9(e)-213.9(it)20.3(s)-215.5(m)0(e)18.6(t)0(a)20.9(b)15(ol)19.3(i)14.3(c)-220.2(a)15(c)0(t)18.8(i)14.3(vi)18.8(ty)-203.8(an)20.5(d)]TJ 0 -1.3858 TD [(to)-331.9(i)14.3(n)0(d)21.6(u)0(c)16.3(e)-339.9(to)20.9(rp)21(or)19.9(.)-342.6(H)0(o)25.4(l)0(e)23.5(s)-347.8(w)15.6(e)0(r)17.8(e)-339.9(dr)21(il)18.7(le)23.5(d)-343(i)14.3(n)-347.2(t)0(h)26.7(e)-346.2(do)24.7(rs)22.5(al)-333.5(c)12.8(a)0(r)19.9(a)0(p)18.4(a)0(c)21.5(e)-339.9(us)21(in)19.9(g)-342(a)]TJ 0 -1.3921 TD [(sy)22(ri)19.3(ng)22.6(e)-258(n)0(e)24.7(e)0(d)16.3(l)0(e)-241.1(a)0(t)-250(p)0(o)24.7(s)0(i)19.3(t)0(i)20.3(o)15(ns)-247.8(on)-244(ei)20.9(th)20.4(e)12.9(r)-259.6(si)25.6(de)-248.2(of)-248.4(th)26.7(e)-258(h)0(e)21(a)0(r)19.9(t)0(,)-242.2(a)0(n)20.5(d)-261.1(th)26.7(in)-251(si)25.6(lv)21.4(er)]TJ 0 -1.3858 TD [(wi)23.6(re)17.8(s)-190.3(\()0(0)22(.)0(2)-184.9(m)0(m)-183.9(d)16.1(ia)16.7(me)24.9(te)18.8(r\))20.4(,)-191.4(i)0(n)19.9(s)0(u)21(l)0(a)25.6(t)0(e)18.8(d)-191.8(ex)21(ce)19.4(pt)-179.6(fo)22.5(r)-196.6(t)0(h)20.4(e)-188.7(ti)20.3(p,)-181.7(we)22.1(re)-183.8(in)19.9(se)24.1(rt)17.2(e)12.8(d)-191.8(by)]TJ 0 -1.3921 TD [(1)12.9(-)0(2)-313.5(m)0(m)-309.8(a)0(n)20.5(d)-324.1(se)24.1(cu)16.3(re)24.1(d)-324.1(i)14.3(n)-322(p)0(l)20.4(a)0(c)21.5(e)0(,)-310.9(a)0(n)20.5(d)-324.1(th)20.4(e)12.9(n)-322(te)18.8(th)26.7(er)17.8(e)12.8(d)-324.1(al)19.3(o)15(n)0(g)-311.2(t)0(h)20.4(e)-321(ex)21(te)18.8(rn)23.1(al)]TJ 0 -1.3858 TD [(su)21(rf)18.8(a)15(c)0(e)-289.3(o)0(f)-292.5(t)0(h)26.7(e)-302.1(c)12.8(a)0(r)19.9(a)0(p)18.4(a)15(ce)19.4(,)-298.5(w)0(i)23.6(t)0(h)-288.3(c)0(y)17.3(a)15(na)20.5(cr)17.8(yl)21.4(a)15(t)0(e)-289.9(a)0(d)18.4(h)14.5(es)17.8(i)14.3(v)0(e)17.3(.)-298.5(Th)23.7(e)-302.1(a)15(ni)19.9(ma)20.7(l)-298(w)0(a)17.9(s)]TJ 0 -1.3921 TD [(th)20.4(e)12.9(n)-315.7(re)24.1(tu)15.7(rn)23.1(e)12.8(d)-317.8(to)-300.4(a)-312.6(h)0(o)23.1(l)0(d)20.4(i)0(n)19.9(g)-316.8(ta)27.2(nk)-305.4(fo)22.5(r)-316.3(2)-314.7(h)14.5(.)-317.4(Du)20.7(ri)25.6(ng)-304.9(th)20.4(is)-302(ti)20.3(me)-302.7(th)20.4(e)-314.7(w)0(i)23.6(r)0(e)17.8(s)]TJ 0 -1.3858 TD [(we)22.1(re)-467.2(co)21.5(nn)17.4(e)12.9(c)12.8(te)18.8(d)-475.3(t)0(o)-470.4(a)-476.4(pu)19.5(rp)21(o)15(s)0(e)17.8(-)13.8(bu)18.4(i)14.3(l)0(t)-468.5(i)0(m)20(p)0(e)22.6(d)0(a)18.4(n)0(c)24.7(e)-478.5(co)21.5(nv)16.3(e)12.8(r)0(t)23.5(e)0(r)-467.2(a)0(n)20.5(d)-481.6(th)26.7(e)]TJ 0 -1.3921 TD [(ou)18.4(tp)22(ut)-198.5(of)-204.3(th)20.4(is)-201.2(wa)17.9(s)-215.5(f)13.8(e)12.8(d)-217(to)-205.9(th)26.7(e)-213.9(i)0(n)19.9(p)0(u)19.5(t)-214.5(o)15(f)-219.2(th)26.7(e)-213.9(P)0(o)16.3(w)15.6(er)17.8(L)14.5(a)0(b)-203.2(i)0(n)19.9(t)0(e)18.8(r)0(f)25.1(a)0(c)21.5(e)0(.)-203.8(I)0(n)-203.2(s)0(o)19.9(m)0(e)]TJ 0 -1.3858 TD [(c)12.9(a)0(s)19.9(e)0(s)17.8(,)-285.9(a)-293.7(w)15.6(in)19.9(do)18.4(w)-286.8(d)0(i)17.8(s)0(c)24.1(r)0(i)19.3(m)0(i)20(n)0(a)26.8(t)0(o)20.9(r)-291.1(fu)23.6(nc)18.4(ti)20.3(o)15(n)-290.5(of)-279.9(th)20.4(e)-289.5(L)14.5(ab)17.3(C)13.5(h)0(a)23.1(r)0(t)-278.9(v)0(7)-278.7(s)0(o)19.9(f)0(t)19.8(w)0(a)24.2(r)0(e)]TJ 0 -1.3921 TD [(wa)24.2(s)-196.6(u)0(s)21(e)12.9(d)-198.1(t)0(o)-180.7(d)0(i)24.1(s)0(p)21(l)0(a)19.3(y)-190.8(th)20.4(e)-195(h)0(e)21(a)0(r)19.9(t)-189.3(ra)19.9(te)-182.8(as)-181.7(a)-192.9(r)0(u)14.7(n)0(n)23.7(i)0(n)19.9(g)-190.8(av)19.4(e)12.8(r)0(a)19.9(g)0(e)-184.3(o)15(f)-194(t)0(h)20.4(e)-195(be)21.5(at)20.9(s)-190.3(p)0(e)16.3(r)]TJ 0 -1.3858 TD [(mi)20(nu)21.6(te)-466.2(\(B)18.9(P)14(M)0(\))18.6(.)-474.9(In)-467.7(o)15(t)0(h)20.4(e)0(r)17.8(s)0(,)-463.7(a)-476.4(ca)21.5(lc)17.2(u)16.1(l)0(a)19.3(t)0(i)20.3(o)15(n)-479.5(o)0(f)-468.8(t)0(h)26.7(e)-478.5(HR)-465.4(wa)17.9(s)-473.8(m)0(a)20.7(d)0(e)-468.7(a)15(t)]TJ 0 -1.3921 TD [(sp)21(e)12.8(c)0(i)]TJ /F7 1 Tf 1.9213 0 TD ()Tj /F6 1 Tf 0.5228 0 TD [(e)12.8(d)-242.2(ti)20.3(me)-227.1(po)18.4(i)14.3(n)0(t)17.8(s)-240.7(b)15(y)-241.2(di)24.1(re)17.8(c)12.9(t)-239.7(co)21.5(un)21.6(ts)-228.5(of)-229.5(th)20.4(e)-239.1(b)0(e)21.5(a)0(t)20.9(s)-240.7(ov)19.4(e)12.8(r)-240.7(a)-237(1)0(5)19.4(s)-240.7(i)14.3(n)0(t)17.8(e)12.9(rv)22(al)19.3(,)]TJ -2.4441 -1.3858 TD [(wh)23.8(ic)20.9(h)-287.9(w)0(e)15.8(r)0(e)-278.3(t)0(h)26.7(e)0(n)-283.9(t)0(r)23.5(a)0(n)20.5(s)0(f)18.8(o)15(rm)23.2(ed)-279.7(to)-281.5(BP)23.7(M.)]TJ 1.5559 -1.3921 TD [(Immediately)-458(prior)-448.7(to)-451.5(electrical)-460.4(stunning)-454.1(the)-451.8(recording)-457(wires)]TJ -1.5559 -1.3858 TD [(were)-218.3(isolated)-220.3(from)-212.5(the)-218.7(electronics)-221.1(and)-215.4(the)-218.7(animals)-218.7(were)-212(subjected)-220.4(to)]TJ 0 -1.3921 TD [(a)-363(standard)-370.1(electrical)-365.9(stun)-370.6(\(5)-362.2(s)-366.7(for)-363.1(crab)-368(and)-366.6(10)-364.8(s)-366.7(for)-363.1(lobster\).)-365(The)]TJ 0 -1.3858 TD 0.0314 Tc [(cardiac)-535.3(r)-1.5(ecording)-531.1(w)2.9(as)-530.6(resumed)-525.8(i)-4.7(mmediately)-531.1(a)-4(fter)-530.6(the)-529(s)-1.5(tun.)]TJ 0 -1.3921 TD 0 Tc [(Ad)-8.7(d)-9.1(i)0(t)-11.2(i)0(o)-8.5(n)0(a)-11(l)0(l)-10.2(y)0(,)-514.5(i)0(n)-515.5(the)-514.8(ca)-10(s)-7.7(e)-510(of)-512.9(s)-7.7(o)0(m)-10.8(e)-510(lob)-16.1(st)-8(ers)-15(,)-512.7(imm)-12.1(ed)-8.9(i)-10.9(ate)-10.3(l)-8.3(y)-505.8(be)-10(fo)-15.3(re)]TJ 0 -1.3858 TD [(stunning)-309.2(one)-306.8(leg)-312.2(was)-304.7(autotomised)-307.9(\(routinely)-310.5(leg)-312.2(L2\))-303.7(and)-309.9(prepared)]TJ T* 0.0378 Tc [(for)-536.8(r)-1.3(ecordin)5.6(g,)-537.9(and)-532(i)-4.6(m)5.7(m)-0.6(ediately)-531(afte)6.6(r)-536.8(s)-1.3(tunning)-537.3(another)-536.8(l)-2(e)6.6(g)]TJ 0 -1.3921 TD 0 Tc [(\(routinely)-360.9(Leg)-352.5(R2\))-353(was)-355.1(also)-357.7(removed)-356.5(and)-354(prepared)-361.1(for)-356.8(recording,)]TJ 0 -1.3858 TD [(before)-303.5(cardiac)-305.9(monitoring)-299.8(was)-304.7(recommenced.)]TJ /F1 1 Tf 11.9999 0 0 11.9999 304.8944 756.6802 Tm [(2.6)-300.5(Ethical)-303.7(statement)]TJ /F6 1 Tf 9 0 0 9 318.8409 731.6786 Tm [(E)-11.2(t)-6.7(hi)-9(ca)-16.3(l)-512.2(a)0(p)-13.1(p)0(r)-10.5(o)0(v)-12.1(a)-10.2(l)-512.2(f)0(o)-9(r)-511.6(p)-9.1(r)-7.7(oc)-10(ed)-15.2(u)-9.1(res)-525.3(on)-514.9(d)-9.1(e)0(c)-12.1(a)0(p)-13.1(o)0(d)-517(c)0(r)-13.7(u)-9.1(s)-7.7(t)-6.7(ac)-10(ea)-16.3(n)-7(s)-511.6(is)]TJ -1.5496 -1.3921 TD [(currently)-387.9(not)-383(required)-390.6(in)-389.5(the)-382.5(UK.)-383.8(Nevertheless,)-396.3(all)-385.8(the)-382.5(live)-386.1(crabs)]TJ 0 -1.3858 TD 0.0142 Tc [(an)-5.4(d)-517.8(l)-0.4(obste)-4.4(rs)-522.6(use)-4.4(d)-517.8(wer)-6.1(e)-514.7(tr)-6.1(eat)-5.1(e)1.9(d)-517.8(w)-1.7(it)-5.1(h)-513.1(p)-1.2(r)-6.1(o)4(p)-7.5(e)1.9(r)-516.3(c)-4.4(are)-521(i)-3(n)-515.7(o)-2.3(rde)-4.4(r)-516.3(to)]TJ 0 -1.3921 TD 0 Tc [(minimise)-489.1(their)-489.2(discomfort)-489.9(and)-492.5(distress.)-494.1(There)-487.3(was)-487.3(no)-489.7(practical)]TJ 0 -1.3858 TD [(alternative)-377.6(to)-369.7(the)-376.2(use)-375.6(of)-374.4(live)-373.5(animals.)-378.6(The)-372.8(total)-375.9(number)-380.8(used)-372.1(\(16)]TJ 0 -1.3921 TD [(crabs)-495.4(and)-492.5(18)-490.8(lobsters\))-496.2(was)-499.9(the)-489.6(minimum)-499.5(required)-497.7(to)-495.6(obtain)-496.2(a)]TJ 0 -1.3858 TD [(rea)-11.3(s)-7.7(o)0(n)-11(a)0(b)-7.9(l)-8.3(e)-510(thr)-12.4(es)-13.7(ho)-8.4(ld)-515(of)-512.9(ce)-12.1(r)-7.7(t)0(a)-10.6(int)-12(y)-8.1(,)-506.4(co)-10(n)-7(s)-7.7(ide)-13.4(r)-7.7(i)0(n)-11.6(g)-505.8(t)-6.7(h)0(a)-8.4(t)-510.6(t)-6.7(h)0(e)-514.4(g)-8.1(ai)-8.5(n)-511(i)0(n)]TJ 0 -1.3921 TD [(knowledge)-312.5(and)-309.9(long-term)-317.9(bene)]TJ /F7 1 Tf 12.5732 0 TD ()Tj /F6 1 Tf 0.5354 0 TD [(t)-309(t)0(o)-313(the)-306.9(subject)-318.6(will)-307.9(be)-312.3(signi)]TJ /F7 1 Tf 11.2252 0 TD ()Tj /F6 1 Tf 0.5354 0 TD (cant.)Tj -24.8693 -1.3858 TD [(Prior)-312.2(to)-313(experimentation)-323.7(they)-315(were)-312.8(housed)-317.5(communally)-314.2(to)-313(ensure)]TJ T* [(behavioural)-353.7(enrichment.)-360.9(Experimental)-359(design)-357.5(considered)-357(the)-357.3(3R)]TJ /F8 1 Tf 26.1984 0 TD ()Tj /F6 1 Tf 0.2205 0 TD (s)Tj -26.4189 -1.3921 TD 0.0315 Tc [(\(Replacem)5.7(ent,)-525.3(Re)6.6(duction)-529.9(a)2.4(nd)-525.7(Re)]TJ /F7 1 Tf 14.7213 0 TD 0 Tc ()Tj /F6 1 Tf 0.5606 0 TD 0.031 Tc [(nemen)5.1(t)-0.9(\).)-532.1(Minimis)4.4(a)-4.4(t)5.4(i)1.2(on)-524.1(of)]TJ -15.2819 -1.3858 TD 0 Tc [(suf)66.3(fering)-384.9(and)-379.2(distress)-384.6(was)-374(inherent)-383.3(to)-382.2(the)-376.2(experimental)-385.3(design)-382.7(in)]TJ 0 -1.3921 TD [(that)-248.5(the)-250.2(objective)-255.2(was)-254.3(to)-250(render)-257.7(the)-250.2(crabs)-256(and)-253.2(lobsters)-253.4(insensible)-258(by)]TJ 0 -1.3858 TD 0.0178 Tc [(e)5.5(l)3.2(ect)4.8(rical)-513.3(s)3.8(tun)4.5(n)-1.8(i)6.9(n)-1.8(g)-513.2(p)2.4(ri)6.9(or)-512.7(to,)-513.8(i)0.6(f)-510.1(n)-1.8(e)5.5(cessary,)-513.8(i)0.6(n)4.5(d)2.4(uc)5.5(in)4.5(g)-519.5(d)2.4(e)5.4(a)1.3(th)7.1(.)-520.1(N)7.2(o)]TJ 0 -1.3921 TD 0 Tc [(anaesthesia)-465(was)-462.1(used.)-462.8(The)-461(exposure)-463.5(of)-462.5(subjects)-464.8(to)-457.8(experimental)]TJ 0 -1.3858 TD [(treatment)-397.7(did)-394.5(not)-395.6(exceed)-396.8(5)-396.6(s)-398.2(for)-394.6(crabs)-394.6(and)-398.1(10)-396.3(s)-391.9(for)-394.6(lobsters.)-400.7(To)]TJ 0 -1.3921 TD [(ensure)-327(welfare,)-326.3(behaviour)-328.9(of)-324(experimental)-328.7(subjects)-326.2(was)-323.6(monitored)]TJ 0 -1.3858 TD [(for)-382(indications)-380.1(of)-386.9(consciousness)-386.7(recovery)-381.2(every)-388.8(15)-377.4(min)-383.8(following)]TJ 0 -1.3921 TD [(stunning.)-330.6(None)-323.7(were)-331.7(apparent)-330.7(up)-326.9(to)-325.6(4)-327.3(h)-325.7(after)-331.7(stunning,)-330.6(when)-329.5(the)]TJ 0 -1.3858 TD [(animals)-300.5(were)-300.2(then)-301.3(euthanised)-304(by)-301.8(freezing.)]TJ /F1 1 Tf 13.5 0 0 13.5 304.8944 456.6613 Tm 0.30196 0.72157 0.33333 scn [(3)-298.9(Results)]TJ 11.9999 0 0 11.9999 304.8944 431.6598 Tm 0.15294 0.15294 0.15294 scn [(3.1)-296.4(The)-303.5(ef)62.9(fectiveness)-304.2(of)-300.3(electrical)-303.7(stunning)]TJ 0 -1.0394 TD [(as)-299.2(a)-302.1(stunning)-304.3(and)-301.9(slaughter)-305.7(procedure)-300.3(for)]TJ /F12 1 Tf 0 -1.0441 TD [(C.)-298.2(maenas)]TJ /F1 1 Tf 10.5001 0 0 10.5001 304.8944 382.1669 Tm [(3.1.1)-296.3(Central)-304.9(nervous)-305.2(system)]TJ /F6 1 Tf 9 0 0 9 318.8409 369.6944 Tm [(Recordings)-391.1(made)-385.1(from)-388.9(the)-382.5(circumoesophageal)-395.5(connectives)-390.4(in)]TJ -1.5496 -1.3921 TD (intact)Tj /F9 1 Tf 2.5575 0 TD [(C.)-298.7(maenas)]TJ /F6 1 Tf 4.4724 0 TD [(crabs)-306.4(\()]TJ 0.30196 0.72157 0.33333 scn 2.7276 0 TD [(Figure)-302.9(1B)]TJ 0.15294 0.15294 0.15294 scn 3.9055 0 TD [(\))-299.5(indicated)-302.8(that)-305.2(there)-301.7(was)-298.4(a)-300(high)]TJ -13.663 -1.3858 TD [(level)-295(of)-298.8(spontaneous)-305.2(neuronal)-305(activity)-300.7(involving)-299.2(a)-300(large)-298.6(number)-298.9(of)]TJ 0 -1.3921 TD 0.0225 Tc [(ax)5.5(ons)-520.6(i)-1(n)-526.3(t)3.2(hese)-525.3(nerves,)-521.7(e)-2.4(ven)-526.3(i)5.3(n)-526.3(t)3.2(he)-525.3(absence)-525.3(of)-524.3(any)-527.4(i)5.3(mposed)]TJ 0 -1.3858 TD 0.0408 Tc [(st)-3.7(im)-3.9(u)-6.1(l)1(a)-7.2(t)2.6(i)-7.9(o)-0.9(n)-545.8(\()]TJ 0.30196 0.72157 0.33333 scn 5.9528 0 TD 0.0454 Tc [(Figu)4.8(re)-540.2(2)]TJ 0.15294 0.15294 0.15294 scn 3.8677 0 TD 0.0458 Tc [(,)-542.5(u)5.2(pper)-541.4(t)7.6(ra)4.1(ces\).)-536.2(In)-540.8(o)4.1(r)0.3(der)-541.4(t)1.3(o)-537.7(t)1.3(est)-540.4(t)7.6(he)]TJ -9.8205 -1.3921 TD 0 Tc [(persistence)-496.5(of)-494(activity)-495.9(in)-496.6(the)-495.9(CNS,)-495.4(some)-497(preparations)-499.4(were)-495.5(re-)]TJ ET 0.63529 0.63529 0.63529 scn 49.606 271.899 m 545.953 271.899 l 545.726 271.672 l 49.833 271.672 l f* 545.953 271.899 m 545.953 70.016 l 545.726 70.242 l 545.726 271.672 l f* 545.953 70.016 m 49.606 70.016 l 49.833 70.242 l 545.726 70.242 l f* 49.606 70.016 m 49.606 271.899 l 49.833 271.672 l 49.833 70.242 l f* q 134.816 259.143 m 460.686 259.143 l 460.686 113.272 l 134.816 113.272 l W n q 325.870789 0 0 145.870834 134.815704 113.272476 cm /Im3 Do Q Q BT /F5 1 Tf 5.5001 0 0 5.5001 60.8315 100.7433 Tm 0.49804 0.49804 0.49412 scn [(FIGURE)-305.5(2)]TJ /F2 1 Tf 6.9999 0 0 6.9999 60.8315 91.7291 Tm [(Recordings)-304.2(of)-305.1(CNS)-301.2(activity)-299.2(in)]TJ /F4 1 Tf 13.4851 0 TD [(C.)-298.4(maenas)]TJ /F2 1 Tf 5.0863 0 TD [(before)-305.1(and)-300.2(after)-304(electrical)-299.8(stunning.)-304.5(Spontaneous)-305.4(nerve)-298.6(activity)-307.3(recorded)-297.3(extracellularly)-305.1(in)-305.7(the)-297.6(left)-305.3(and)-300.2(right)]TJ -18.5713 -1.2878 TD [(circumoesophageal)-313.3(connectives)-298.4(\(COC\))-306.3(\(arrows\).)-302.1(Upper)-303.9(traces)-302.1(intact)-305.8(crab;)-296.6(Lower)-305.3(traces:)-302.7(10)-302(min)-298.7(after)-304(electrical)-299.9(stunning.)-304.5(Scale)-299.8(bar,)-306.4(1)-295.2(s)0(.)]TJ /F1 1 Tf -1.6036 102.6079 TD [(Neil)-304.8(et)-292.7(al.)]TJ 57.4715 0 TD (10.3389/fanim.2024.1378350)Tj -57.4715 -109.3302 TD [(Frontiers)-304.6(in)]TJ 0.30196 0.72157 0.33333 scn 5.5074 0 TD [(Animal)-299.2(Science)]TJ 58.8564 0 TD (frontiersin.org)Tj 0.15294 0.15294 0.15294 scn -29.5213 0 TD (06)Tj ET endstream endobj 119 0 obj<>stream HyLSՊD&q(Q T*$jĘh9R px ZQTR,EhBp 'p-xiEghsYIZAAAAAAAAAAA|OK+     TVVȴ4Bڨ{+W U*n544D"@ Je\=!!^|911UUUi~B0===)))k׮AC]U*/^B,y&.~EgY 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[(nerve)-306.7(\(4)-300.4(V)-301.2(a)0(t)-299(various)-301(frequencies\).)-308.9(Scale)-299.8(bars,)-298.7(3)-298.4(s)-308.1(and)-300.2(500)-295.7(mN.)]TJ ET 0.63529 0.63529 0.63529 scn 49.606 765.524 m 545.953 765.524 l 545.726 765.297 l 49.833 765.297 l f* 545.953 765.524 m 545.953 550.091 l 545.726 550.318 l 545.726 765.297 l f* 545.953 550.091 m 49.606 550.091 l 49.833 550.318 l 545.726 550.318 l f* 49.606 550.091 m 49.606 765.524 l 49.833 765.297 l 49.833 550.318 l f* q 134.816 752.825 m 460.686 752.825 l 460.686 602.362 l 134.816 602.362 l W n q 325.870789 0 0 150.462967 134.815704 602.362122 cm /Im5 Do Q Q BT /F5 1 Tf 5.5001 0 0 5.5001 60.8315 589.8329 Tm 0.49804 0.49804 0.49412 scn [(FIGURE)-305.5(3)]TJ /F2 1 Tf 6.9999 0 0 6.9999 60.8315 580.8188 Tm [(Recordings)-304.2(of)-305.1(sensory)-300.2(activity)-307.3(in)]TJ /F4 1 Tf 14.951 0 TD [(C.)-298.4(maenas)]TJ /F2 1 Tf 5.0863 0 TD [(before)-305.1(and)-300.2(after)-304(electrical)-299.8(stunning.)-304.5(Responses)-306.9(of)-297(the)-305.7(leg)-297.3(nerve)-306.7(to)-300.3(three)-298.8(forms)-300.9(of)-305.1(stimulation)-301(of)-297(the)]TJ -20.0373 -1.2878 TD [(dactylopodite,)-312.9(as)-300.3(indicated.)-299.4(Upper)-303.9(trace,)-297.7(leg)-305.4(R2)-303.3(autotomised)-299.5(from)-300.5(intact)-305.8(animal)-303.2(before)-297(stunning;)-309.2(lower)-301(trace,)-297.7(leg)-305.4(L2)-304.9(amputated)-298.7(from)-300.5(same)-300.2(animal)]TJ 0 -1.2796 TD [(after)-304(electrical)-307.9(stunning.)-304.5(Scale)-299.8(bar,)-298.3(10)-302(s.)]TJ /F1 1 Tf -1.6036 34.0164 TD [(Neil)-304.8(et)-292.7(al.)]TJ 57.4715 0 TD (10.3389/fanim.2024.1378350)Tj -57.4715 -109.3302 TD [(Frontiers)-304.6(in)]TJ 0.30196 0.72157 0.33333 scn 5.5074 0 TD [(Animal)-299.2(Science)]TJ 58.8564 0 TD (frontiersin.org)Tj 0.15294 0.15294 0.15294 scn -29.4809 0 TD (07)Tj ET endstream endobj 132 0 obj<>stream H}HU)67Uwq6_ RZ(hT*FYG&&JJZhI򨭹pis 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[(Spontaneous)-305.4(activity)-307.3(recorded)-297.3(in)-305.7(the)-297.6(left)-305.3(circumoesophageal)]TJ -39.8801 -1.2797 TD (connective.)Tj /F5 1 Tf 5.556 0 TD (\(B\))Tj /F2 1 Tf 1.5388 0 TD [(Spontaneous)-305.4(activity)-299.2(recorded)-305.4(from)-300.5(the)-297.6(left)-305.3(connective)-298.1(between)-306.1(segments)-301.3(2)-304.6(and)-300.2(3)-298.4(o)0(f)-305.1(the)-297.6(abdomen.)]TJ /F5 1 Tf 45.8978 0 TD (\(C\))Tj /F2 1 Tf 1.6117 0 TD [(Responses)-298.8(of)-305.1(the)-297.6(leg)-305.5(nerve)]TJ -54.6044 -1.2878 TD [(to)-308.4(three)-298.8(forms)-300.9(of)-297(stimulation)-309.1(of)-297(the)-305.7(dactylopodite,)-304.8(as)-300.3(indicated.)-307.5(Top)-293.4(trace,)-305.8(leg)-305.4(R2)-295.2(autotomised)-307.6(from)-300.5(intact)-305.8(animal)-295.1(before)-305.1(stunning;)-301.1(lower)-301(trace,)-305.8(leg)]TJ T* [(L2)-304.9(amputated)-298.7(from)-308.6(same)-300.2(animal)-303.2(after)-295.9(electrical)-307.9(stunning.)]TJ /F5 1 Tf 26.2493 0 TD (\(D\))Tj /F2 1 Tf 1.6279 0 TD [(Spontaneous)-305.4(activity)-307.3(recorded)-297.3(in)-305.7(the)-297.6(3)]TJ 5.2498 0 0 4.6661 378.7653 569.6503 Tm (rd)Tj 6.9999 0 0 6.9999 385.7952 566.5322 Tm [(motor)-305.3(root)-296.1(of)-305.1(the)-297.6(fourth)-306.8(abdominal)-305.1(segment.)]TJ -46.4242 -1.2878 TD [(In)-305.2(all)-299.8(panels,)-307.3(upper)-301.3(traces:)-302.7(animal)-303.2(before)-297(electrical)-307.9(stunning;)-301.1(lower)-301(traces:)-302.7(animal)-303.2(10)-302(min)-298.7(after)-304(electrical)-299.8(stunning.)-304.5(Scale)-299.8(bars,)-306.8(1)-303.3(s)-300(in)]TJ /F5 1 Tf 59.3747 0 TD [(\(A,)-301.4(B,)-303.7(D\))]TJ /F2 1 Tf 4.0253 0 TD [(10)-302(s)]TJ -63.4 -1.2797 TD (in)Tj /F5 1 Tf 1.1177 0 TD (\(C\))Tj /F2 1 Tf 1.3121 0 TD (.)Tj /F1 1 Tf -4.0334 36.0573 TD [(Neil)-304.8(et)-292.7(al.)]TJ 57.4715 0 TD (10.3389/fanim.2024.1378350)Tj -57.4715 -109.3302 TD [(Frontiers)-304.6(in)]TJ 0.30196 0.72157 0.33333 scn 5.5074 0 TD [(Animal)-299.2(Science)]TJ 58.8564 0 TD (frontiersin.org)Tj 0.15294 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[(senti)-4.4(e)5.8(n)-21.4(c)20.4(e)-253.3(a)-5.6(nd)-246.5(s)-5.7(u)4.6(s)-5.7(t)5.2(a)-5.6(i)3.7(n)-21.4(a)17.1(ble)-253.3(s)2.4(ea)-5.6(food.)]TJ /F9 1 Tf 13.1611 0 TD -0.0176 Tc [(Na)9.7(t.)-250.3(F)8.4(o)-17.6(o)19.4(d)]TJ /F6 1 Tf 4.0091 0 TD -0.0254 Tc [(3,)-260.9(88)-8(4)]TJ /F8 1 Tf 2.2759 0 TD 0 Tc ()Tj /F6 1 Tf 0.494 0 TD -0.025 Tc [(88)-7.6(6.)-260.6(doi)-8(:)]TJ 0.15294 0.15294 0.15294 scn 3.4826 0 TD -0.0187 Tc [(10)6.7(.1)6.7(03)6.8(8/s)5.1(4)-1.4(3)6.7(0)-1.4(1)6.8(6)-1.4(-02)6.8(2)-1.4(-0)6.8(06)6.7(32)6.7(-6)]TJ 0.13725 0.12157 0.12549 scn -22.4913 -1.5712 TD 0 Tc [(DEFRA)-224.9(\(2021\))-224.7(Our)-231.7(action)-224.9(plan)-228.2(for)-221.9(animal)-227.4(welfare.)-227.4(Available)-233.2(online)-221.5(at:)]TJ 0.30196 0.72157 0.33333 scn 28.1526 0 TD (https://assets.)Tj -29.084 -1.142 TD [(pu)-12.9(b)-10.2(l)-9.2(ish)-19.4(in)-17(g.)-12.4(se)-13.7(rv)-12.2(ic)-12.4(e)-13.3(.)0(g)-12.4(o)-10.2(v.)-12.4(uk)-11.5(/)-10.1(g)0(o)-13.9(v)0(e)-16.9(r)0(n)-10.2(m)0(e)-19.3(n)0(t)-15.5(/up)-23(lo)-11.3(ad)-14.9(s/)-10.5(sy)-20.3(st)-14.3(em)-11.2(/)-10.1(u)0(p)-12.9(l)-9.2(oa)-10.6(ds)-15(/)-10.1(a)0(t)-14.2(t)0(a)-14.2(c)0(h)-16.8(m)0(e)-11.2(n)-9.7(t_d)-22.5(at)-14.2(a/)]TJ /F7 1 Tf 32.8987 0 TD ()Tj /F6 1 Tf 0.5426 0 TD [(le)-14.3(/)]TJ -33.4413 -1.142 TD (985332/Action_Plan_for_Animal_Welfare.pdf)Tj 0.13725 0.12157 0.12549 scn 18.5389 0 TD (.)Tj -17.6075 -1.5712 TD [(DEFRA)-386.9(\(2022\))-378.5(Animal)-390.4(welfare)-388.7(\(sentience\))-388.2(act)-383.9(2022)-384.2(\(UK)-384(Government\).)-390.7(Available)]TJ -0.9314 -1.142 TD [(online)-302.5(at:)]TJ 0.30196 0.72157 0.33333 scn 4.0658 0 TD (https://bills.parliament.uk/bills/2867)Tj 0.13725 0.12157 0.12549 scn 14.4894 0 TD (.)Tj -17.6237 -1.5712 TD [(DeFur,)-302.6(P.)-313.4(L.,)-304.6(and)-308.1(Mangum,)-308.3(C.)-308.5(P.)-305.3(\(1979\).)-306.3(The)-309.9(ef)59.1(fects)-308.7(of)-302.4(environmental)-309.4(variables)-312.6(on)]TJ -0.9314 -1.1501 TD [(the)-314.2(heart)-323.1(rates)-320(of)-318.6(invertebrates.)]TJ /F9 1 Tf 12.9262 0 TD [(Comp.)-317.6(Biochem.)-320.1(Physiol.)]TJ /F6 1 Tf 10.1806 0 TD [(62A,)-315.7(283)]TJ /F8 1 Tf 3.6203 0 TD ()Tj /F6 1 Tf 0.5102 0 TD [(294.)-313.1(doi:)]TJ 6.9999 0 0 6.9999 266.7968 449.348 Tm 0.15294 0.15294 0.15294 scn (10.1016/)Tj -31.0278 -1.142 TD (0300-9629\(79\)90058-6)Tj 0.13725 0.12157 0.12549 scn 0.9314 -1.5712 TD [(EFSA)-330.5(\(2005\).)-330.6(Aspects)-340.8(of)-326.7(the)-338.5(biology)-329.9(and)-332.4(welfare)-332(of)-334.8(animals)-333.1(used)-334.4(for)-327.2(experimental)]TJ -0.9314 -1.142 TD [(and)-283.8(other)-276.3(scienti)]TJ /F7 1 Tf 6.7304 0 TD ()Tj /F6 1 Tf 0.5345 0 TD [(c)-280.5(purposes.)]TJ /F9 1 Tf 4.7623 0 TD [(Asp.)-279.2(Biol.)-279.2(Welf.)-285(Anim.)-278.7(Used)-278(Exp.)-280.6(other)-277.4(Sci.)-279.8(Purp)]TJ /F6 1 Tf 19.4946 0 TD [(292,)-272.6(1)]TJ /F8 1 Tf 2.4135 0 TD ()Tj /F6 1 Tf -33.9354 -1.142 TD [(46.)-298.1(doi:)]TJ 0.15294 0.15294 0.15294 scn 3.2801 0 TD (10.2903/j.efsa.2005.292)Tj 0.13725 0.12157 0.12549 scn -2.3487 -1.5712 TD [(El)-14.1(wo)-13.6(o)-10.2(d)0(,)-517.4(R)-9.6(.)-519(W)0(.)-522.9(\()0(2)-12.2(0)0(1)-13.9(9)-15(\).)-516.2(Di)-23.4(sc)-13.7(ri)-15.8(mi)-13.3(n)-9.7(a)0(t)-14.2(i)0(o)-17.5(n)-511.9(b)-10.2(e)0(t)-10.9(w)-11.4(ee)-18.4(n)-511.9(n)-9.7(o)-10.2(c)0(i)-12.4(c)0(e)-18.4(p)0(t)-12.2(i)0(v)-19(e)-515.4(re)]TJ /F7 1 Tf 26.6219 0 TD ()Tj /F6 1 Tf 0.5426 0 TD [(ex)-15(es)-524(an)-10.1(d)-516.7(m)0(o)-16.3(r)0(e)]TJ -28.0959 -1.142 TD [(complex)-432.8(responses)-427.4(consistent)-432.4(with)-433(pain)-428.8(in)-430.1(crustaceans.)]TJ /F9 1 Tf 23.3741 0 TD [(Phil.)-435.4(Trans.)-425.4(R.)-433.4(Soc)-424.1(B.)]TJ /F6 1 Tf 9.4112 0 TD (374,)Tj -32.7853 -1.142 TD [(20190368.)-299.3(doi:)]TJ 0.15294 0.15294 0.15294 scn 6.1553 0 TD (10.1098/rstb.2019.0368)Tj 0.13725 0.12157 0.12549 scn -5.2239 -1.5712 TD [(Factor,)-482.7(J.)-480.5(R.)-480.1(\(1995\).)]TJ /F9 1 Tf 8.8766 0 TD [(The)-478.2(biology)-487(of)-476.9(lobsters)]TJ /F6 1 Tf 9.8485 0 TD [(\(San)-486.9(Diego)-479.9(USA:)-481.7(Academic)-482.5(Press\).)]TJ -19.6566 -1.142 TD (doi:)Tj 0.15294 0.15294 0.15294 scn 1.798 0 TD (10.1016/B978-0-12-247570-2.X5021-X)Tj 0.13725 0.12157 0.12549 scn -0.8666 -1.5712 TD [(FAO)-240.8(\(2020\).)]TJ /F8 1 Tf 5.2482 0 TD ()Tj /F6 1 Tf 0.3969 0 TD [(The)-237(state)-244.3(of)-237.6(world)]TJ /F7 1 Tf 7.4431 0 TD ()Tj /F6 1 Tf 0.5264 0 TD [(sheries)-249.4(and)-235.2(aquaculture)-248.6(2020,)]TJ /F8 1 Tf 11.7761 0 TD ()Tj /F6 1 Tf 0.6317 0 TD (in)Tj /F9 1 Tf 1.0367 0 TD [(Sustainability)-253.3(in)]TJ -27.9906 -1.142 TD (action)Tj /F6 1 Tf 2.4378 0 TD [(\(Rome:)-304.3(FAO\).)-303.5(doi:)]TJ 0.15294 0.15294 0.15294 scn 7.8157 0 TD (10.4060/ca9229en)Tj 0.13725 0.12157 0.12549 scn -9.3221 -1.5712 TD [(FAO)-378.5(\(2021\).)-379.2(Fishstat)-386.5(data)-377.4(\(Rome:)-385.3(FAO\).)-384.5(Available)-379(at:)]TJ 0.30196 0.72157 0.33333 scn 22.6533 0 TD (http://www.fao.org/)Tj /F7 1 Tf 7.8643 0 TD ()Tj /F6 1 Tf 0.5345 0 TD (shery/)Tj -31.9835 -1.1501 TD (statistics/en2021)Tj 0.13725 0.12157 0.12549 scn 6.5765 0 TD (.)Tj -5.6451 -1.2797 TD -0.0356 Tc [(Fregin,)-214.5(T)-4.7(.,)-222.6(and)-220.2(B)-3.3(i)5.7(c)-0.3(kmey)-6.9(er,)-222.6(U)5(.)-222.6(\()-0.4(201)]TJ 12.5537 0 TD [(6\).)-222.5(E)0.1(lectrophy)-6.8(s)4.5(iological)-214.8(i)-2.3(nv)-6.8(es)]TJ 10.5451 0 TD -0.0344 Tc [(tigat)8.4(i)-1.2(on)-214.2(of)-221.2(dif)62.2(f)5.5(erent)-218.4(m)0.1(ethods)]TJ -24.0301 -1.142 TD -0.0351 Tc [(of)-302.9(anesthesia)-302.7(in)-295.9(lobster)-294.7(a)-3.1(nd)-300.7(cray)]TJ /F7 1 Tf 12.0434 0 TD 0 Tc ()Tj /F6 1 Tf 0.494 0 TD -0.0319 Tc [(sh)5.1(.)]TJ /F9 1 Tf 1.304 0 TD -0.0336 Tc [(PloS)-296.4(One)]TJ /F6 1 Tf 3.6932 0 TD [(11,)-301.4(1)]TJ /F8 1 Tf 1.8142 0 TD 0 Tc ()Tj /F6 1 Tf 0.4778 0 TD -0.0346 Tc [(19)7(.)-302.5(d)-0.6(oi:)]TJ 0.15294 0.15294 0.15294 scn 3.0291 0 TD -0.034 Tc [(1)7.6(0)-0.5(.13)7.6(71/journ)4.9(a)-2(l)5.4(.)-2.3(pon)4.8(e)1.3(.016)7.6(2894)]TJ 0.13725 0.12157 0.12549 scn -21.9244 -1.5712 TD 0.3237 Tc [(G)351.4(r)355.6(e)350.9(e)350.9(n)354.5(,J)354.1(.A)354.7(.\()350.8(2)349.1(0)357.2(1)349.2(1)349.1(\))350.8(.)7.2(T)346.5(h)352.6(eh)352.6(e)350.9(a)355.7(r)347.5(t)2.1(r)355.6(a)347.6(t)350.3(e)2.7(m)350.1(e)350.9(t)358.4(h)352.6(o)345.9(d)1.4(f)355.5(o)]TJ 17.6885 0 TD -0.0253 Tc [(r)-349.7(e)1.9(st)9.4(ima)6.7(t)1.3(in)5.5(g)-344.8(m)1.1(eta)6.7(b)5(ol)6(ic)-346.3(r)6.6(a)-1.4(te)10(:)-349.9(a)-349.6(r)6.6(e)1.9(view)-344.5(an)5.5(d)]TJ -18.6199 -1.1501 TD -0.0282 Tc [(reco)-6.1(m)6.3(m)-1.8(endat)6.5(i)-3.1(ons.)]TJ /F9 1 Tf 7.2325 0 TD -0.0269 Tc [(Comp.)-267.6(B)2.5(ioch)5.3(em.)-259.5(P)-1.3(hy)5.1(siol.)]TJ /F6 1 Tf 9.4355 0 TD -0.0254 Tc [(1)8.1(58A)5.6(,)-261(287)]TJ /F8 1 Tf 3.8147 0 TD 0 Tc ()Tj /F6 1 Tf 0.4859 0 TD -0.0261 Tc [(3)7.4(04.)-261.6(doi:)]TJ 0.15294 0.15294 0.15294 scn 3.4583 0 TD -0.0244 Tc [(10)9.1(.1)9.2(016)9.1(/j)6.9(.)7.3(c)2.8(bp)9.7(a.)7.3(201)9.2(0.)7.3(09.)7.3(0)1.1(1)9.1(1)]TJ 0.13725 0.12157 0.12549 scn -23.4956 -1.5712 TD 0 Tc [(Gregory,)-490.8(N.)-500.9(G.,)-492.1(and)-494.4(Wotton,)-491.9(S.)-496.5(B.)-494.9(\(1987\).)-492.6(Ef)59.3(fect)-494.5(of)-496.8(electrical)-498.4(stunning)-498.4(on)-489.8(the)]TJ -0.9314 -1.142 TD [(electroencephalogram)-220.2(in)-211.4(chickens.)]TJ /F9 1 Tf 13.8657 0 TD [(Br.)-211.9(Vet.)-214.1(J.)]TJ /F6 1 Tf 3.9281 0 TD [(143,)-215.9(175)]TJ /F8 1 Tf 3.3125 0 TD ()Tj /F6 1 Tf 0.5102 0 TD [(183.)-207.8(doi:)]TJ 0.15294 0.15294 0.15294 scn 3.5717 0 TD (10.1016/0007-1935\(87\))Tj -25.1883 -1.142 TD (90009-1)Tj 0.13725 0.12157 0.12549 scn 0.9314 -1.5712 TD (Grif)Tj /F7 1 Tf 1.5631 0 TD ()Tj /F6 1 Tf 0.5264 0 TD [(n,)-229.1(R.)-229(A.,)-221.5(McCuaig,)-232.1(J.)-229.4(D.,)-228.1(and)-227.1(Blewett,)-221.5(T.)-229(A.)-228.9(\(2023\).)]TJ /F8 1 Tf 19.9887 0 TD ()Tj /F6 1 Tf 0.3969 0 TD [(Ecotoxicology)-233.4(and)-227.1(response)]TJ -23.4065 -1.142 TD [(to)-421(pollutants,)]TJ /F8 1 Tf 5.4264 0 TD ()Tj /F6 1 Tf 0.818 0 TD (in)Tj /F9 1 Tf 1.223 0 TD [(Ecophysiology)-420(of)-420.2(the)-420.9(european)-422(green)-426(crab)-423.1(\()]TJ /F6 1 Tf 17.7209 0 TD [(Carcinus)-418.8(maenas)]TJ /F9 1 Tf 6.9896 0 TD [(\))-423.1(and)]TJ /F6 1 Tf -32.1779 -1.142 TD [(related)-267.4(species.)-265.7(Eds.)-263.6(D.)-267.9(Weihrauch)-270.8(and)-259.5(I.)-267(J.)-261.8(McGaw)-270.6(\(London)-265.9(UK)-257.2(and)-267.6(San)-262.9(Diego)-269.3(USA:)]TJ T* [(Elsevier)-303.6(Inc\).)-303.4(doi:)]TJ 0.15294 0.15294 0.15294 scn 7.3459 0 TD (10.1016/B978-0-323-99694-5.00001-5)Tj 0.13725 0.12157 0.12549 scn -6.4145 -1.5712 TD 0.0241 Tc [(Han)6.3(d)-6.6(y,)-527.3(R.)-519.2(D.,)-527.3(a)-0.5(nd)-525(Dep)-6.6(l)-1.3(edge,)-527.3(M)2(.)-527.3(H)-1.5(.)-519.2(\()-5.5(1999\))-5.5(.)-527.3(P)3(hysiologic)-5.4(al)-519.6(re)-5.3(spon)6.3(se)-5.4(s:)-519.2(Their)]TJ -0.9314 -1.142 TD 0 Tc [(measurement)-415.3(and)-413.4(use)-417(as)-413.9(environmental)-414.7(biomarkers)-418.4(in)-413.9(ecotoxicology.)]TJ /F9 1 Tf 29.1326 0 TD (Ecotoxicology)Tj /F6 1 Tf -29.1326 -1.142 TD [(8,)-299.2(329)]TJ /F8 1 Tf 2.4378 0 TD ()Tj /F6 1 Tf 0.5102 0 TD [(349.)-296.9(doi:)]TJ 0.15294 0.15294 0.15294 scn 3.758 0 TD (10.1023/A:1008930404461)Tj 0.13725 0.12157 0.12549 scn -5.7747 -1.5712 TD 0.5595 Tc [(H)533.9(a)534.9(r)526.6(r)534.7(i)527.9(n)533.6(g)531.5(t)529.4(o)533.1(n)533.6(,A)533.8(.M)537.4(.)526.4(,)8.1(H)525.8(a)534.8(v)531.6(e)530(r)534.7(k)530.2(a)526.8(m)537.3(p)528.8(,)0(H)533.9(.)534.5(,a)534.9(n)533.6(d)2.3(H)525.8(a)534.9(m)529.2(l)534.1(i)527.9(n)533.6(,)8.1(H)525.8(.)8.1(J)525.1(.)8.1(\()529.9(2)528.3(0)536.4(2)528.3(0)536.3(\))529.9(.)0(I)536.1(m)529.2(p)528.8(e)538.1(d)528.8(a)526.8(n)533.6(c)530(e)]TJ -0.9314 -1.142 TD 0 Tc [(p)-14.5(n)-9.7(e)-13.3(u)-14.5(m)-14.1(o)-18.3(g)-11.8(r)0(a)-25.1(p)-14.5(h)-11.6(y)-530.1(fo)-27(r)-526.9(m)-14.1(in)-25.1(i)-15.4(m)0(a)-22.6(l)-17.3(l)-9.2(y)-530.1(in)-25.1(v)-11.8(a)-16.6(si)-23.9(v)-11.7(e)-531.6(m)-14.1(e)-13.3(as)-25.1(u)-14.5(r)0(e)-21.8(m)-14.1(e)-13.3(n)-17.8(t)-524.1(o)-10.2(f)-527(h)-19.7(e)-13.3(a)0(r)-25.1(t)-524.1(r)-16.7(a)0(t)-22.3(e)-531.6(in)-535.4(l)-17.3(a)0(t)-22.3(e)-531.6(st)-22.4(a)-16.6(g)-11.7(e)]TJ 0 -1.1501 TD (invertebrates.)Tj /F9 1 Tf 5.6613 0 TD [(J.)-301.3(Vis.)-308.1(Exp.)]TJ /F6 1 Tf 4.6732 0 TD [(158\),)-302.2(e61096.)-299.8(doi:)]TJ 0.15294 0.15294 0.15294 scn 7.435 0 TD (10.3791/61096)Tj 0.13725 0.12157 0.12549 scn -16.8381 -1.5712 TD [(Hjelmstedt,)-354.9(P.,)-362.7(Sundell,)-356.4(E.,)-362.7(Brijs,)-358.4(J.,)-351.6(Berg,)-366.5(C.,)-349.7(Sandblom,)-362.5(E.,)-362.7(Lines,)-359(J.,)-351.6(et)-359.2(al.)-358.5(\(2022\).)]TJ -0.9314 -1.142 TD [(Assessing)-247.9(the)-249.4(ef)59.1(fectiveness)-251.1(of)-245.7(percussive)-259.4(and)-243.3(electrical)-255.4(stunning)-247.3(in)-251.9(rainbow)-244.2(trout:)-248.1(Does)]TJ T* [(an)-366.5(epileptic-like)-378.6(seizure)-370.3(imply)-372.8(brain)-368.3(failure?)]TJ /F9 1 Tf 18.4093 0 TD (Aquaculture)Tj /F6 1 Tf 5.2644 0 TD [(552,)-369.8(738012.)-366.4(doi:)]TJ 0.15294 0.15294 0.15294 scn 7.354 0 TD (10.1016/)Tj -31.0278 -1.142 TD (j.aquaculture.2022.738012)Tj 0.13725 0.12157 0.12549 scn 0.9314 -1.5712 TD [(Ho)28.9(r)15.7(r)15.7(i)0(d)26.8(g)0(e)23.6(,)-211.3(G)19.6(.)-211.3(A.)22.2(,)-203.2(a)0(n)22.3(d)-208.9(Sa)30.3(nd)24.3(em)29.3(an)22.3(,)-211.3(D)16.3(.)-211.3(C)16.2(.)-211.3(\()0(1)28.3(9)17.3(64)26.6(\).)-200.3(N)18.2(e)0(r)26.8(v)0(o)26.6(u)0(s)-193.2(c)0(o)25.1(n)0(t)25(r)15.7(ol)-197.6(of)-197.1(op)31.9(to)24.5(k)19.2(i)0(n)23.5(e)0(t)29.6(i)17(c)-215.7(r)15.7(es)26.8(po)31.9(ns)22.2(es)]TJ 6.9999 0 0 6.9999 49.6063 124.3842 Tm [(i)17(n)-293.2(t)18.5(h)0(e)-276(c)19.1(ra)23.5(b)]TJ /F9 1 Tf 4.5436 0 TD -0.0176 Tc [(Ca)-17.6(r)9.3(c)-17.6(i)11.9(n)-17.6(u)7.7(s)-0.5(.)]TJ /F6 1 Tf 3.92 0 TD 0 Tc [(Pr)27(oc)25.1(.)]TJ /F9 1 Tf 2.2921 0 TD -0.0201 Tc [(R.)-309.4(S)-20.1(o)6.9(c)-301.1(\()-20.1(B\))]TJ /F6 1 Tf 4.1387 0 TD 0 Tc [(1)17.3(6)0(1)26.6(,)-284.1(21)26.6(6)]TJ /F8 1 Tf 3.2963 0 TD ()Tj /F6 1 Tf 0.494 0 TD -0.0172 Tc [(24)-17.2(6)9.4(.)-301.3(d)-17.2(o)6.6(i:)]TJ 0.15294 0.15294 0.15294 scn 3.6446 0 TD 0 Tc [(1)17.4(0)17.3(.1)24.8(0)17.4(9)0(8)26.6(/)0(r)21.9(s)15.7(p)17.9(b)0(.)21.5(1)17.4(9)17.3(6)0(4)26.6(.)15.5(00)26.6(91)]TJ 0.13725 0.12157 0.12549 scn -21.3979 -1.5712 TD [(Hvingel,)-486.2(C.,)-495.5(Sainte-Marie,)-489.7(B.,)-487.5(and)-494.4(Kruse,)-489.2(G.)-491.4(H.)-488(\(2021\).)-492.6(Cold-water)-488.9(shell)]TJ /F7 1 Tf 30.7929 0 TD ()Tj /F6 1 Tf 0.5345 0 TD [(sh)-481.9(as)]TJ -32.2588 -1.142 TD [(harvestable)-279.3(resources)-277.1(and)-283.8(important)-278.9(ecosystem)-277(players.)]TJ /F9 1 Tf 22.4832 0 TD [(ICES)-275.5(J.)-277(Mar.)-278.4(Sci.)]TJ /F6 1 Tf 6.9005 0 TD [(78,)-281.9(479)]TJ /F8 1 Tf 2.8914 0 TD ()Tj /F6 1 Tf 0.5103 0 TD (490.)Tj -32.7853 -1.142 TD (doi:)Tj 0.15294 0.15294 0.15294 scn 1.798 0 TD (10.1093/icesjms/fsab005)Tj 0.13725 0.12157 0.12549 scn -0.8666 -1.5712 TD [(Kells,)-241.5(N.)-249.8(J.,)-246.3(Perrott,)-252.2(M.,)-242.2(and)-251.4(Johnson,)-249.2(C.)-243.7(B.)-243.8(\(2023\).)-249.6(The)-245.1(ef)]TJ /F7 1 Tf 22.6614 0 TD ()Tj /F6 1 Tf 0.5345 0 TD [(cacy)-241.1(of)-245.7(electrical)-255.4(stunning)]TJ -24.1273 -1.142 TD [(of)-310.5(New)-314.3(Zealand)-313(rock)-312.5(lobster)-308.8(\()]TJ /F9 1 Tf 12.1811 0 TD [(Jasus)-318.4(edwardsii)]TJ /F6 1 Tf 6.1067 0 TD [(\))-304.9(and)-316.2(freshwater)-316.9(cray)]TJ /F7 1 Tf 8.6175 0 TD ()Tj /F6 1 Tf 0.5264 0 TD [(sh)-311.8(\()]TJ /F9 1 Tf 1.555 0 TD (Paranephrops)Tj 7.4836 97.4245 TD [(ze)-15.1(a)-13.1(l)0(a)-17.1(n)0(d)-13.8(i)-13.5(cu)-17.3(s)]TJ /F6 1 Tf 4.6651 0 TD [(\))-523.6(u)0(s)-15(i)0(n)-17(g)-522(th)-17.4(e)-515.4(C)0(r)-24.8(u)0(s)-15(t)-13.9(as)-17(tu)-20.3(n)]TJ /F10 1 Tf 6.5001 0 0 6.5001 406.2613 761.8959 Tm ()Tj /F6 1 Tf 6.9999 0 0 6.9999 412.6676 760.3652 Tm (.)Tj /F9 1 Tf 0.737 0 TD [(An)-19.8(im)-18.1(.)-516.1(W)0(e)-18.7(l)-12.1(fa)-18.5(re)]TJ /F6 1 Tf 6.698 0 TD [(32)-13.9(,)-519(1)]TJ /F8 1 Tf 2.203 0 TD ()Tj /F6 1 Tf 0.5183 0 TD [(9,)-526(e5)-20.2(7.)-526(do)-16.7(i:)]TJ 0.15294 0.15294 0.15294 scn 5.4021 0 TD 0.0151 Tc [(10)15.1(.1)15.1(017)15.1(/)]TJ -30.9549 -1.142 TD 0 Tc (awf.2023.76)Tj 0.13725 0.12157 0.12549 scn 0.9233 -1.5712 TD [(Kholodkevich,)-441(S.)-439.8(V.,)-438.3(Kuznetsova,)-442.8(T.)-439.6(V.,)-438.3(Sharov,)-441.9(A.)-439.5(N.,)-436.8(Kurakin,)-441.3(A.)-439.5(S.,)-440.5(Lips,)-439.6(U.,)]TJ -0.9233 -1.142 TD [(Kolesova,)-305.5(N.,)-315.3(et)-302.5(al.)-309.9(\(2017\).)-314.4(Applicability)-314.7(of)-310.5(a)-308.1(bioelectronic)-309.9(cardiac)-308.7(monitoring)-313.5(system)]TJ T* [(for)-270.5(the)-273.7(detection)-279.5(of)-270(biological)-275.4(ef)59.1(fects)-276.3(of)-270(pollution)-277.1(in)-268.1(bioindicator)-283.1(species)-273.2(in)-276.2(the)-273.7(Gulf)-272(of)]TJ T* (Finland.)Tj /F9 1 Tf 3.5879 0 TD [(J.)-301.3(Mar.)-302.7(Syst.)]TJ /F6 1 Tf 5.1591 0 TD [(171,)-305(151)]TJ /F8 1 Tf 3.3935 0 TD ()Tj /F6 1 Tf 0.5103 0 TD [(158.)-305(doi:)]TJ 0.15294 0.15294 0.15294 scn 3.758 0 TD (10.1016/j.jmarsys.2016.12.005)Tj 0.13725 0.12157 0.12549 scn -15.4855 -1.5712 TD [(Kl)25.1(a)15.8(s)15.7(se)26.8(n,)-318.2(G.)27(,)-340.8(a)15.8(nd)-315.9(Lo)26.7(ck)30.3(e,)-321.7(A.)-318(\(2)20.2(0)17.3(0)17.4(7\))28.3(.)-340.8(A)-333.5(bi)31.1(ol)29.2(og)26.6(i)17(c)0(a)26.9(l)-333.1(s)15.7(y)0(n)27.1(o)0(p)23.8(s)15.7(i)17(s)-340.6(o)22.2(f)-340.7(t)18.5(h)0(e)-324.6(e)19.1(ur)25.5(op)31.9(ea)26.9(n)-333.7(g)0(r)28.3(e)0(e)22.1(n)-333.7(c)19.1(r)0(a)23.5(b)0(,)]TJ /F9 1 Tf -0.9233 -1.142 TD [(c)18.6(a)0(r)26.9(c)0(i)29.4(n)0(u)25.3(s)-306.8(ma)30.8(en)27(as)]TJ /F6 1 Tf 6.4064 0 TD (.)Tj /F9 1 Tf 0.5264 0 TD [(C)17.6(a)0(n)23.8(.)-305.5(M)16.5(a)0(n)23.8(u)0(s)29.8(c)0(r)26.3(.)-305.5(Re)24.2(p)17.3(.)-305.5(Fi)28.7(sh)25(.)-305.5(A)0(q)25.4(u)0(a)31.9(t)0(.)-295.1(S)0(c)31.2(i)]TJ /F6 1 Tf 14.457 0 TD [(no)28.6(.)-308.4(2)17.3(8)17.3(1)0(8)26.6(:)-308.4(vi)29.6(i+)22.1(7)17.3(5)17.3(p)0(p)19.5(.)-300.3(Fi)26.8(sh)28.4(er)26.8(ie)28.1(s)-308.2(a)0(n)22.3(d)]TJ -21.3898 -1.1501 TD -0.0222 Tc [(Oc)-22.2(e)8(a)-22.2(ns)-306.1(C)-6(a)-22.2(na)-6.4(da)-22.2(,)-298.6(M)-22.2(on)-22.2(c)11.5(t)-22.2(on)-22.2(,)-291.8(N)-22.2(B.)]TJ 0.9233 -1.5712 TD 0 Tc [(Kushinsky,)-221.8(D.,)-220(Morozova,)-217.1(E.)-216.2(O.,)-218.1(and)-210.9(Marder,)-222(E.)-216.2(\(2019\).)]TJ /F9 1 Tf 22.0701 0 TD [(In)-213.2(vivo)]TJ /F6 1 Tf 2.9076 0 TD [(ef)59.1(fects)-219.6(of)-213.3(temperature)]TJ -25.901 -1.142 TD [(on)-311.6(the)-322.3(heart)-323.1(and)-316.2(pyloric)-325.9(rhythms)-315.1(in)-324.8(the)-314.2(crab)]TJ /F9 1 Tf 18.6118 0 TD [(Cancer)-323(borealis)]TJ /F6 1 Tf 6.1391 0 TD (.)Tj /F9 1 Tf 0.5345 0 TD [(J.)-325.6(Exp.)-313(Biol.)]TJ /F6 1 Tf 5.0134 0 TD [(222,)-313.1(1)]TJ /F8 1 Tf 2.454 0 TD ()Tj /F6 1 Tf 0.5102 0 TD (12.)Tj -33.2631 -1.142 TD (doi:)Tj 0.15294 0.15294 0.15294 scn 1.798 0 TD (10.1242/jeb.199190)Tj 0.13725 0.12157 0.12549 scn -0.8747 -1.5712 TD [(Lowe,)-338.9(S.,)-343.3(Browne,)-337.6(M.,)-339.4(Boudjelas,)-348(S.,)-335.2(and)-340.5(De)-337.2(Poorter,)-345.8(M.)-338.7(\(2000\).)]TJ /F9 1 Tf 26.3627 0 TD [(100)-345.4(of)-339.2(the)-340(world)]TJ /F15 1 Tf 6.6089 0 TD ()Tj /F9 1 Tf 0.2106 0 TD (s)Tj -34.1054 -1.142 TD [(worst)-240.1(invasive)-247.8(alien)-246.6(species:)-244.4(A)-243(selection)-247.6(from)-247.1(the)-242.8(global)-245.6(invasive)-247.8(species)-238.6(database)]TJ /F6 1 Tf 31.5704 0 TD [(Vol.)-246.4(12)]TJ -31.5704 -1.142 TD [(\(Auckland:)-300.9(Invasive)-304.8(Species)-306.8(Specialist)-300(Group\).)]TJ 0.9233 -1.5712 TD [(Mar)-14.9(der)-12.1(,)-502.8(E.,)-516.6(an)-10.1(d)-500.5(Buc)-11.8(her,)-520.1(D.)-510.9(\(20)-11.1(07\))-11.1(.)-502.8(U)0(n)-9.7(der)-12.1(stan)-16.4(ding)-513.1(ci)-12.4(rcui)-11.3(t)-507.9(dyn)-11.8(amic)-10.7(s)-510.7(usin)-15.9(g)-505.8(the)]TJ -0.9233 -1.142 TD [(stomatogastric)-329.4(nervous)-335.9(system)-337.4(of)-334.8(lobsters)-333.6(and)-332.4(crabs.)]TJ /F9 1 Tf 21.6814 0 TD [(Ann.)-337(Rev.)-330.3(Physiol.)]TJ /F6 1 Tf 7.6456 0 TD [(69,)-330.5(291)]TJ /F8 1 Tf 2.94 0 TD ()Tj /F6 1 Tf 0.5102 0 TD (316.)Tj -32.7772 -1.142 TD (doi:)Tj 0.15294 0.15294 0.15294 scn 1.798 0 TD (10.1146/annurev.physiol.69.031905.161516)Tj 0.13725 0.12157 0.12549 scn -0.8747 -1.5712 TD [(Marom,)-388.2(S.,)-391.9(and)-389.1(Marder,)-392.1(E.)-386.3(\(2023\).)-387.3(A)-390.2(biophysical)-387.9(perspective)-391.4(on)-392.6(the)-387(resilience)-387.2(of)]TJ -0.9233 -1.142 TD [(neuronal)-310.6(excitability)-311.9(across)-308.8(timescales.)]TJ /F9 1 Tf 15.8419 0 TD [(Nat.)-315.9(Rev.)-306(Neurosci.)]TJ /F6 1 Tf 7.9696 0 TD [(24,)-314.3(640)]TJ /F8 1 Tf 2.9238 0 TD ()Tj /F6 1 Tf 0.5102 0 TD [(652.)-313.1(doi:)]TJ 0.15294 0.15294 0.15294 scn 3.7742 0 TD (10.1038/)Tj -31.0197 -1.1501 TD (s41583-023-00730-9)Tj 0.13725 0.12157 0.12549 scn 0.9233 -1.5712 TD [(McGaw,)-425.2(I.)-420.9(J.,)-416.4(and)-421.5(Reiber,)-419.3(C.)-421.9(L.)-417.3(\(2015\).)]TJ /F8 1 Tf 16.749 0 TD ()Tj /F6 1 Tf 0.3888 0 TD [(Circulatory)-429.8(physiology,)]TJ /F8 1 Tf 9.476 0 TD ()Tj /F6 1 Tf 0.8099 0 TD (in)Tj /F9 1 Tf 1.223 0 TD [(The)-421.5(natural)]TJ -29.5699 -1.142 TD [(history)-272.3(of)-266.3(crustaceans)]TJ /F6 1 Tf 8.4798 0 TD [(,)-276(vol.)-266.8(4)-266.1(.)-276(Eds.)-271.7(E.)-264.8(S.)-269.7(Chang)-276.6(and)-267.6(M.)-273.9(Thiel)-269.7(\(Oxford)-273.9(University)-270.2(Press,)]TJ -8.4798 -1.142 TD [(USA\),)-300.7(199)]TJ /F8 1 Tf 4.1792 0 TD ()Tj /F6 1 Tf 0.5102 0 TD (246.)Tj -3.7661 -1.5712 TD [(McLean,)-257.4(K.)-258(M.,)-258.4(and)-251.4(Todgham,)-259.3(A.)-253.2(E.)-256.7(\(2015\).)-257.7(Ef)59.3(fect)-251.6(of)-197.1(food)-254.3(availability)-259(on)-254.9(the)-249.4(growth)]TJ -0.9233 -1.142 TD [(and)-243.3(thermal)-249.2(physiology)-244.7(of)-245.7(juvenile)-250.2(Dungeness)-251.4(crabs)-243.5(\()]TJ /F9 1 Tf 21.3169 0 TD [(Metacarcinus)-243.3(magister)]TJ /F6 1 Tf 8.9252 0 TD (\).)Tj /F9 1 Tf 0.8099 0 TD (Conserv.)Tj -31.0521 -1.142 TD (Physiol.)Tj /F6 1 Tf 3.3368 0 TD [(3,)-299.2(cov013.)-307.9(doi:)]TJ 0.15294 0.15294 0.15294 scn 6.131 0 TD (10.1093/conphys/cov013)Tj 0.13725 0.12157 0.12549 scn -8.5446 -1.5712 TD [(McVean,)-237.2(A.)-228.9(\(1975\).)-233.5(Autotomy:)-228.5(mini)-232.8(review.)]TJ /F9 1 Tf 17.7047 0 TD [(Comp.)-228.5(Biochem.)-231(Physiol.)-234.1(A)]TJ /F6 1 Tf 10.7961 0 TD [(51,)-233.3(497)]TJ /F8 1 Tf 2.8428 0 TD ()Tj /F6 1 Tf 0.5183 0 TD (505.)Tj -32.7853 -1.142 TD (doi:)Tj 0.15294 0.15294 0.15294 scn 1.798 0 TD (10.1016/0300-9629\(75\)90332-1)Tj 0.13725 0.12157 0.12549 scn -0.8747 -1.5712 TD [(Neil,)-295.7(D.)-300.3(M.,)-290.8(Albalat,)-296.4(A.,)-302.5(and)-291.9(Thompson,)-300.1(J.)-294.2(\(2022\).)-298.2(The)-293.7(ef)59.1(fects)-300.6(of)-294.3(electrical)-295.9(stunning)]TJ -0.9233 -1.142 TD [(on)-271.1(the)-281.8(nervous)-279.2(activity)-281.9(and)-275.7(physiological)-279.3(stress)-280.1(response)-281.2(of)-278.1(a)-283.8(commercially)-278.5(important)]TJ T* [(de)-11.6(ca)-13.6(pod)-525.3(cr)-13.7(us)-15(ta)-14.2(ce)-10.3(an)-18.2(,)-510.9(t)0(h)-17.4(e)-507.3(b)-10.2(r)0(o)-10.7(w)-11.4(n)-511.9(c)0(r)-13.7(a)0(b)]TJ /F9 1 Tf 16.0444 0 TD [(Ca)-11.8(nc)-9.2(e)-10(r)-510.7(pa)-12(g)-13.3(a)0(r)-13.6(u)0(s)]TJ /F6 1 Tf 6.9815 0 TD (L.)Tj /F9 1 Tf 1.2635 0 TD [(Pl)-18.9(oS)-515.7(On)-11.2(e)]TJ /F6 1 Tf 4.4059 0 TD [(17,)-524.8(e0)-12.1(27)-13.9(09)-13.9(60)-13.9(.)]TJ -28.6952 -1.1501 TD (doi:)Tj 0.15294 0.15294 0.15294 scn 1.798 0 TD (10.1371/journal.pone.0270960)Tj 0.13725 0.12157 0.12549 scn -0.8747 -1.5712 TD [(Passantino,)-503.7(A.,)-488.8(Elwood,)-498.7(R.)-496.3(W.,)-491.2(and)-502.5(Coluccio,)-491.9(P.)-499.7(\(2021\).)-492.6(Why)-497.1(protect)-492.6(decapod)]TJ -0.9233 -1.142 TD [(crustaceans)-501.3(used)-504.5(as)-503(models)-507.3(in)-494.9(biom)-9.5(edical)-503.4(research)-506.8(and)-502.5(in)-494.9(ecotoxicology?)]TJ /F9 1 Tf 31.6919 0 TD (Ethical)Tj -31.6919 -1.142 TD [(Legislative)-304.7(Considerat)-299.1(Anim.)]TJ /F6 1 Tf 11.7275 0 TD [(11,)-306.2(73.)-298.1(doi:)]TJ 0.15294 0.15294 0.15294 scn 4.7542 0 TD (10.3390/ani11010073)Tj 0.13725 0.12157 0.12549 scn -15.5584 -1.5712 TD [(Phillips,)-308.2(B.)-308.6(F.)-306.8(\(2013\).)]TJ /F9 1 Tf 8.8038 0 TD [(Lobsters:)-307.9(biology,)-306.6(management,)-306.3(aquaculture)-312.8(and)]TJ /F16 1 Tf 19.5594 0 TD ()Tj /F9 1 Tf 0.5264 0 TD (sheries)Tj /F6 1 Tf 2.5998 0 TD (.)Tj /F9 1 Tf 0.5265 0 TD (2nd)Tj 6.9999 0 0 6.9999 304.8944 428.3716 Tm (ed)Tj /F6 1 Tf 1.1906 0 TD [(\(Oxford)-306.3(UK:)-298.4(Wiley\).)-302.4(doi:)]TJ 0.15294 0.15294 0.15294 scn 10.4641 0 TD (10.1002/9781118517444)Tj 0.13725 0.12157 0.12549 scn -10.7313 -1.5712 TD [(Powell,)-245.2(D.)-235.5(J.,)-246.3(Owens,)-236.1(E.,)-241.2(Bergsund,)-252(M.)-233.4(M.,)-242.2(Cooper,)-243.9(M.,)-242.2(Newstein,)-246.3(P.,)-241.2(Berner,)-240.5(E.,)-241.2(et)-237.7(al.)]TJ -0.9233 -1.142 TD 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Neil Endre Putyora Amaya Albalat Neil DM, Putyora E and Albalat A (2024) Towards the humane slaughter of decapod crustaceans: identifying the most effective indicators of insensibility following electrical stunning. Front. Anim. Sci. 5:1378350 Copyright © 2024 Neil, Putyora and Albalat. This is an open-access article distributed under the terms of the Creative Commons Attribution License (CC BY). The use, distribution or reproduction in other forums is permitted, provided the original author(s) and the copyright owner(s) are credited and that the original publication in this journal is cited, in accordance with accepted academic practice. No use, distribution or reproduction is permitted which does not comply with these terms. electrical stunning humane slaughter decapod crustaceans insensibility cardiac activity Towards the humane slaughter of decapod crustaceans: identifying the most effective indicators of insensibility following electrical stunning electrical stunning, humane slaughter, decapod crustaceans, insensibility, cardiac activity True https://www.frontiersin.org/legal/copyright-statement endstream endobj xref 0 1 0000000469 65535 f 469 2 0000000000 00001 f 0002323687 00000 n 472 1 0002323819 00000 n 716 1 0002324022 00000 n trailer << /Size 717 /Prev 116 /Root 472 0 R /ID [ <4E40AE6D0E2D6F0C58BD1E020C61BC16> <7f460377A8EBC342BD0C28C9ED68FE6E> ] /Info 470 0 R >> %EndExifToolUpdate 2323665 startxref 2328913 %%EOF 682 0 obj <> endobj 683 0 obj <> endobj 693 0 obj <> endobj 696 0 obj <> endobj 697 0 obj <> endobj 699 0 obj <> endobj 703 0 obj <> endobj 716 0 obj <>stream application/pdf Douglas M. Neil Endre Putyora Amaya Albalat Decapod crustaceans provide a valuable food source worldwide, both through wild capture and captive rearing. Copyright © 2024 Neil, Putyora and Albalat. This is an open-access article distributed under the terms of the Creative Commons Attribution License (CC BY). The use, distribution or reproduction in other forums is permitted, provided the original author(s) and the copyright owner(s) are credited and that the original publication in this journal is cited, in accordance with accepted academic practice. No use, distribution or reproduction is permitted which does not comply with these terms. 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